The Science of Replenishment

Anaplerosis

The metabolic pathways that replenish the TCA cycle hold the key to treating some of medicine's most devastating inherited metabolic disorders.

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4
TCA Cycle Entry Points
~1:50K
Anaplerotic Disorder Prevalence
2020
FDA Triheptanoin Approval

The Science of Metabolic Replenishment

Anaplerosis—from the Greek ana (up) and plerotikos (to fill)—describes the biochemical reactions that replenish intermediates of the tricarboxylic acid (TCA) cycle. When these pathways fail due to inherited enzyme deficiencies, the consequences cascade through every energy-dependent tissue in the body: brain, heart, liver, and skeletal muscle.

The 2020 FDA approval of triheptanoin (Dojolvi®) for long-chain fatty acid oxidation disorders marked a watershed moment—the first therapy designed explicitly to exploit anaplerotic biochemistry. But this is just the beginning. Research into anaplerotic substrates, gene therapy for TCA cycle enzymes, and metabolic flux analysis is reshaping how we approach inborn errors of metabolism.

This resource explores the science, the therapies, and the clinical genetics behind anaplerotic medicine—authored by board-certified clinical geneticists who manage these conditions every day.

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